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Francis, an intelligent and cynical cat, is moved to a gloomy, dilapidated house with his owner Gustav Löbel, an architect and erotic novelist. The top floor of the house gives off a mysterious and distinctly chemical odor. During Francis's exploratory tour of the house, he discovers the body of a cat who was killed by a bite to the neck. At the crime scene, he meets and befriends Bluebeard, a foul-mouthed, one-eyed and mutilated Maine Coon. Soon, Francis discovers another body and the fact that his house's top floor is the meeting site of a sect of cats led by Joker; the members of the sect worship an entity named Claudandus and perform ritualistic suicide by throwing themselves in an electric current. When Francis's cover is blown, he is chased by the sect members through the city's rooftops. He escapes through a skylight and meets a blind cat named Felicity, who supplies Francis with information on the Claudandus sect. The next day, Bluebeard takes Francis to Pascal, an elderly and tech-savvy cat who has taken to meticulously maintaining a list of feline deaths in the area, though which Francis learns that Felicity is the latest victim. That night, Francis is haunted by a nightmare in which the famous friar and geneticist Gregor Mendel leaves him with riddles. During a rodent hunt, Francis discovers a video recording that documents the top floor's previous use as an experimental laboratory; this laboratory was devoted to the research and development of a fibrin glue and tissue adhesive that would close wounds in an instant. The test subjects largely consisted of stray cats that, more often than not, died in agony as a result of the failed test runs. The one cat who survived the experiments was christened by the lab's technicians as "Claudandus", Latin for "He who should or must be sealed". However, Claudandus eventually murdered the project's lead technician Dr. Preterius, leading to the escape of the other strays and the closure of the lab. Claudandus would subsequently become a legendary martyr figure revered by the modern-day sect...

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